This article relates to the official games. See Pikipedia's "Dusk Pustules" article for more official information.

Dusk Pustule

Fruit

Appearance in Pikmin 4.

Dusk Pustules are a fruit, specifically a bunch of black muscat grapes. In Pikmin 3, where it acts as a fruit, Pikmin pluck individual Dawn Pustules from the bunch until no more are left, and in Pikmin 4, where it acts as a treasure, Pikmin carry the entire bunch whole.

In Pikmin Bloom, Dusk Pustules are giant fruits that yield blue nectar.

Appearance in Pikmin 3 ().
Appearance in Pikmin Bloom ().

In fanon games

Below this point is where users place their version of the Dusk Pustule.

None yet!

Official fruits

Astringent Clump • Blonde Impostor • Citrus Lump • Combustion Berry • Crimson Banquet • Crunchy Deluge • Cupid's Grenade • Dapper Blob • Dawn Pustule • Delectable Bouquet • Disguised Delicacy • Dusk Pustule • Face Wrinkler • Fire-Breathing Feast • Heroine's Tear • Indigo Morsel • Insect Condo • Juicy Gaggle • Lesser Mock Bottom • Mock Bottom • Pocked Airhead • Portable Sunset • Scaly Custard • Searing Acidshock • Seed Hive • Slapstick Crescent • Special fruit • Stellar Extrusion • Sunseed Berry • Tremendous Sniffer • Velvety Dreamdrop • Wayward Moon • Zest Bomb

Pikmin: Wide World

Locations
Planets	Main (PNF-404) • Side (Hocotate) • Mentioned (Koppai)
Areas	Balmy Highlands • Arid Oasis • Frigid Tundra • Ancient Metalworks • Colossal Hollow • Sprawling Swamp
Dens and caves	Temperate Retreat • Grassy Grotto • Dune Den • Sunken Sands • Desolate Ditch • Humid Burrow • Snowy Recess • Lonely Quarry • Watering Hole • Warm Sett • Flaming Trench • Sunbaked Sauna • Gritty Getaway • Wasted Waterworks • Dim Greenhouse • Clogged Pipeline • Wooden Fortress • Stone Dugout • Icy Crevasse • Hidden Scrapyard • Frozen Lodge • Castaway Cove • Concrete Compound • Chemical Reservoir • Subterranean Spring • Dreary Sepulcher • Fungal Jungle • Dank Depths • Forgotten Mine • Wayward Citadel • Lost Kingdom • Polluted Pond • Corroded Complex • Robot Workshop • Sunless Sanctuary • Watery Grave • Overgrown Garden • Muddy Murk • Buzzing Reserve • Webbed Lair • Secluded Hideaway • Divers' Blight • Sunlit Pit • Black Abyss • Pest Nest • Neglected Aqueduct • Cursed Chasm • Destiny Cavern
Battle Mode stages	Close Quarters • Forest Frenzy • Tan Battleground • Cold War • Planked Peril • Hard Fought • Reinforced Fortress • Frantic Factory • Tiled Tossup • Mashup Mayhem
Challenge Mode stages	Pretty Prairie • Dingy Dugout • Forgotten Garden • Serene Shore • The Colossal Cottonade • Weathered Field • Arctic Abode • Preserved Pit • Frozen Fields • The Brumal Arcticlops • Rocky Road • Cement City • Rebar Labyrinth • Steel Stronghold • The Plasm Wraith • Sandstone Citadel • Perplexing Paradise • Corroded Catwalk • Jellyfloat Pool • The Titan Weaver Dweevil • Impenetrable Prison • Foreboding Forest • Timber Maze • Mandiblard Nest • The Hydroe Host • Study Floor • Wooden Expanse • Mosaic Maze • Breadbug Den • The Grimacing Cremblub

Characters

Main (Olimar • Shacho • Marie • Louie • Dolphin Lander) • Side (Alph • Brittany • Charlie • Dorris • Hunnigan • Granny Marma • Libra • Margot • Sagittarius • S.S. Drake • S.S. Dreamstride) • Mentioned (Bulbie • Freight Voyager • SPERO)

Pikmin

Main (Yellow Pikmin • Blue Pikmin • Red Pikmin • Rock Pikmin • Green Pikmin • Purple Pikmin • White Pikmin • Winged Pikmin) • Other (Alien Pikmin • Captmin • Mushroom Pikmin) • Parasitic Pikmin (Blue Bulbmin • Breadmin • Burrowing Snagmin • Cannon Beetlemin • Dweevilmin • Groinkmin • Hairy Bulbmin • Orange Bulbmin • Red Bulbmin • Spinermin • Spotty Bulbearmin • Wollymin) • Utilities (Onion • Pellet)

Piklopedia

Enemies (?)

Acid Dumple • Acidic Jellyfloat • Adult Green Centipare • Adult Red Centipare • Agitated Flint Beetle • Alloy Spectralid • Ancient Bulblax • Angler Bulblax • Anodestool • Aquatic Cannon Grub • Arctic Cannon Grub • Ardent Dirigibug • Armored Cannon Beetle • Armored Cannon Grub • Armurk • Baldy Long Legs • Ballistic Groink • Balloon Mitite • Battering Groink • Beady Long Legs • Bearded Amprat • Berry Bloyster • Bilious Bulbot • Black Spectralid • Bladed Greater Beeb • Blazing Cannon Grub • Blubbug • Blue Bulborb • Bombardier Beeb • Bomber Dweevil • Breadbug • Briary Burrow-nit • Brilliant Glowstalk • Brumal Arcticlops • Bubble-Blowing Blowhog • Bug-Eyed Crawmad • Bulbear Larva • Bulborb Larva • Bullhop • Bumbling Snitchbug • Buoyant Blubbug • Burrow-nit Larva • Burrowing Snagret • Buttery Breadbug • Calcified Crushblat • Carbonated Goolix • Careening Dirigibug • Carnation Bulborb • Carnivorous Candypop • Cave Bulbat • Ceiling Slurker • Ceramic Joustmite • Chaotic Kettlebug • Charred Skitterleaf • Chemical Bulblax • Chill Dirigibug • Cinnamon Breadbug • Clawed Long Legs • Clicking Slurker • Cloaking Burrow-nit • Clod Dirigibug • Clumsy Blubbug • Colossal Cottonade • Common Kettlebug • Coppeller • Coriaceous Elephog • Corpulent Breadblax • Corrosion Beetle • Crammed Hermer • Crawbster Young • Cream Spectralid • Creeping Chrysanthemum • Crested Mockiwi • Cryoplasmic Slooch • Crystalline Crushblat • Daddy Long Legs • Dangling Splurchin • Decorated Cannon Beetle • Decorated Cannon Grub • Deranging Bloyster • Desiccated Skitterleaf • Diamond Long Legs • Dousing Dweevil • Dwarf Blue Bulborb • Dwarf Electric Bulbeel • Dwarf Searing Bulblax • Dwarf Hairy Bulborb • Dwarf Orange Bulborb • Dwarf Puckering Bulbeel • Dwarf Red Bulborb • Dwarf Spotty Bulbear • Dwarf Stubborn Bulblon • Dwarf Vampire Bulbear • Dwarf Vitreous Bulborb • Dwarf Whiptongue Bulborb • Dwarf Yellow Bulborb • Electric Bulbeel • Electric Cottonade • Electric Spectralid • Electric Starnacle • Electripede • Elongated Bulborb • Emberglow Phosbat • Emperor Bulblax • Emperor Bulbot • Empress Orange Bulblax • Empress Red Bulblax • Empress Spotty Bulblax • Fandangled Isleclops • Female Sheargrub • Female Shooting Spiner • Ferrying Coppeller • Fiendish Flytrap • Fiery Wollyhop • Firebreathing Blowhog • Firebreathing Blowlet • Fireflap Bulborb • Fireflinger Groink • Flaming Mitite • Flatterchuck • Flatulent Flint Beetle • Flittering Glowbelly • Floating Oobuoy • Flourishing Blowhog • Fluttery Splurchin • Flying Joustmite • Foraging Cottonade • Foundation Dweevil • Fountain Crawmad • Framewing Spectralid • Freezing Skeeterskate • Frosted Breadbug • Frosthop Blowhog • Frosty Bulblax • Fuzzy Blubbug • Galvanic Jellyfloat • Gambler Clamclamp • Gatling Groink • Gaudy Crawmad • Gelidity Dweevil • Gemstone Kettlebug • Ghost Wolpole • Ghost Wollyhop • Giant Breadbug • Giant Buttery Breadbug • Giant Cinnamon Breadbug • Giant Frosted Breadbug • Giant Ginger Breadbug • Giant Licorice Breadbug • Gilded Flint Beetle • Gilled Mawdad • Ginger Breadbug • Glassed Wollyhop • Golden Clamclamp • Goolix • Grabbit • Greasy Jellyfloat • Greater Bubblimp • Greater Jellyfloat • Green Bulbot • Green Centipare • Grimacing Cremblub • Grouchy Goblax • Hailstorm Dandelfly • Hairy Bulborb • Harvester Mamuta • Headdress Slurker • Heir Bulblax • Hermit Crawmad • Honey Bubblimp • Honeypot Skeeterskate • Honeywisp • Hopping Clamclamp • Hopping Flint Beetle • Humming Kettlebug • Humped Bloyster • Hungry Starnacle • Hydroe Host • Ignition Dweevil • Immaculate Whiskerfly • Incandescent Beeb • Incendiary Spectralid • Inky Waddlepus • Iridescent Flint Beetle • Ivory Waddlepus • Jamboree Blowhog • Joustmite • Juggling Dirigibug • Juicy Capstool • King Mawdad • Large Splurchin • Leech Hydroe • Lesser Bubblimp • Lesser Jellyfloat • Licorice Breadbug • Little Long Legs • Loafling • Lofty Mockiwi • Long Dumple • Luminous Long Legs • Lurking Lotus • Maggot Whiskerpillar • Magnetic Joustmite • Male Sheargrub • Male Shooting Spiner • Mamuta • Man-at-Legs • Medusal Slurker • Mephitic Mitite • Mesmerizing Sputtlefish • Metal Shearwig • Mighty Grabbit • Miniature Mudleech • Mitite • Mockiwi • Monochromatic Pinchipede • Monster Dumple • Moon Jellyfloat • Mossy Long Legs • Mouthy Mimitator • Muddy Meerslug • Mudleech • Mudwisp • Muggonfly • Mung Dweevil • Naked Amprat • Nectar Goolix • Nectarous Dandelfly • Needlesnout Blowhog • Neon Blinnow • Noxious Crawbster • Oily Groink • Orange Bulborb • Orebagger • Overcharge Blowhog • Painted Whiskerpillar • Pale Wolpole • Pale Wollyhop • Peacap Puffstalk • Peach Bulborb • Pearl Sparrowhead • Pearly Clamclamp • Peckish Aristocrab • Pellet Beetle • Pellet Poser • Phosbat Nymph • Pileated Snagret • Pinching Sheargrub • Plain Spectralid • Plasm Wraith • Plated Kettlebug • Plump Sheargrub • Poison Dart Wollyhop • Praying Skittermantis • Prickly Splurchin • Prodigious Beeb • Puckering Blinnow • Puckering Bulbeel • Puffstalk • Puffstool • Puffy Blowhog • Puffy Blowlet • Putrid Bulbear • Pygmy Jellyfloat • Pyroclasmic Slooch • Quaggled Mireclops • Queen Shearwig • Raging Long Legs • Rainwisp • Ranging Bloyster • Ravenous Loccut • Red Bulborb • Red Bulbot • Red Centipare • Red Spectralid • Reef Blowhog • Regurgitating Twistool • Ringed Spiner • Robust Bulbot • Rocket Beetle • Rocky Hermer • Ruby Jellyfloat • Rusted Cannon Beetle • Sable Flint Flicker • Sandbelching Meerslug • Sandy Speargrub • Saplapper Mockiwi • Satchel Snitchbug • Scornet • Scornet Maestro • Searchlight Snitchbug • Searing Bulblax • Seedbagger • Segmented Crawbster • Shady Sparrowhead • Shaggy Long Legs • Shearblug • Shearpion • Shearwig • Shelled Meerslug • Shock Dumple • Shore Dweevil • Shrimpy Hydroe • Silent Glowbelly • Skeeterskate • Skitterflake • Skitterleaf • Skitterlion • Skitterreed • Skittish Piddlepoddler • Skutterchuck • Sky Dumple • Sleepy Bulbear • Slick Dweevil • Slithering Shallock • Smoky Progg • Snaglet • Soaring Jellyfloat • Sparkling Dirigibug • Speargrub • Spectral Slurker • Spiked Snagret • Spinose Mockiwi • Spiny Coppeller • Spitty Blowhog • Spitty Blowlet • Sporegrub • Spornet • Spotty Bulbear • Spouting Bellbloom • Sprinting Dirigibug • Sprouting Coppeller • Sputtersplash • Sputtlefish • Stamping Long Legs • Starnacle • Steely Flint Beetle • Stingwig • Stubborn Bulblon • Studded Greater Beeb • Stuffed Bellbloom • Sulfuric Dweevil • Surge Dweevil • Swarming Sheargrub • Swooping Snitchbug • Tainting Blowhog • Tan Sparrowhead • Tantrum Snagret • Telescoping Pumphog • Titan Dweevil • Titan Weaver Dweevil • Toady Bloyster • Tombcarver Mawdad • Trampling Mamuta • Translucent Wollybot • Tropical Bloyster • Twilight Spectralid • Twinkle Beetle • Vampire Bulbear • Vapor Goolix • Vehemoth Phosbat • Venomous Scornet • Virescent Pinchipede • Vitreous Bulborb • Volatile Splurchin • Volt Beetle • Voracious Whiskerpillar • Waddlepus • Water Dumple • Water Wraith • Weaver Dweevil • Web-Slinging Spiner • Webbed Mockiwi • Whiptongue Bulborb • White Spectralid • Widemouthed Corrosion Beetle • Widemouthed Volt Beetle • Winged Snagret • Winter Snownet • Withering Blowhog • Withering Blowlet • Wolfish Mitite • Woolly Mammhog • Worker Beeb • Wracknid Hatchling • Wracknid Mother • Yellow Bulborb • Yellow Bulbot • Yellow Spectralid • Yellow Wolpole • Yellow Wollybot • Yellow Wollyhop • Young Fiery Wollyhop • Young Ghost Wollyhop • Young Pale Wollyhop • Young Yellow Wollyhop

Vegetation

Asphyxiating Moldclump • Bloominous Stemple • Bouncy Mushroom • Bramble • Browncap • Burgeoning Spiderwort • Capstalk • Carnation Candypop • Colossal Glowcap • Common Glowcap • Crimson Candypop • Demantoid Candypop • Flukeweed • Golden Candypop • Hydroe Bramble • Ivory Candypop • Kingcap • Lapis Lazuli Candypop • Lilypad • Monarch Candypop • Mouthy Shootplant • Nectar Grass • Orbular Pollenspout • Pellet Posy • Pellet Vine • Shockcap • Slate Candypop • Sparklium Candypop • Spiny Spiderwort • Spotcap • Springpetal • Sunset Spiderwort • Thorned Posy • Vine • Violet Candypop • Weeping Posy • Wickweed

Collectibles

Data file • Fruit (list) • Nugget • Sparklium • Treasure • Upgrades
Black nectar • Hammer seed • Knuckle seed • Nectar • Ultra-bitter spray • Ultra-hot spray • Ultra-minty spray • Ultra-spicy spray • Ultra-sweet spray • Ultra-toxic spray • Whip seed

Hazards

Acid • Adhesive • Blunt force • Bubble • Electricity • Explosion • Fire • Ice • Lubricant • Mud • Panic • Poison • Spore • Water

Obstacles and objects

Acid pool • Bamboo gate • Bomb rock • Boulder • Brick gate • Burrow-nit nest • Cardboard box • Climbing stick • Clipboard • Clog • Cobblestone block • Conveyor belt • Conveyor switch • Cryogenic bomb rock • Crystal • Crystal gate • Crystal nodule • Crystal stalactite • Dazzling bomb rock • Egg • Electric bulb gate • Electric wire gate • Electrical wire • Electrode • Elevator block • Elevator platform • Escape geyser • Fire geyser • Fire rhizome gate • Flame • Flukeweed dam • Fragment bridge (bridge fragment) • Frozen surface • Gas pipe • Glowstem • Hay slope (hay stack) • Ice chunk • Incendiary bomb rock • Iron ball • Jackwig hive • Liftable elevator • Metal block • Metal pipe • Mine • Mine rock • Mitite bomb • Mud puddle • Nectar pool • Oil slick • Phosbat pod • Pit • Plasm • Poison rhizome gate • Pulley elevator • Rain droplet • Rectangular paper bag • Reinforced gate • Rhizome gate • Rock • Rubble • Rusted rod • Sand pit • Sky Dumple cluster • Smoke bomb rock • Snowball • Soil block • Soil gate • Soil mound • Stick bridge • Sweetgum pod • Tin box • Triangular paper bag • Trowel • Tunnel • Water geyser • Water pool • Web • Wooden stake

Gameplay

Achievements • Caravan • Dialogue • Hints • HocoPad • Murals • Options • Pikpik carrot • Soul • Soundtrack • Squad desertion • Sunset • Temperature • Weather • Game modes (Story Mode • Battle Mode • Challenge Mode • Bottomless Abyss • Randomizer)

Other

Changes during development • Gallery